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Jack takes Sophie home, but afterward a mysterious female voice lures him to Pitch's lair, where Tooth's fairies and the teeth are being held.

Although slightly complicated by the intrusion of Jamie's sister, Sophie, the preparation goes smoothly. The dejected Guardians rally by helping Bunnymund prepare his eggs for Easter. Despite Jack's efforts to save him, Sandy is overwhelmed and seemingly killed by Pitch. Pitch learns of the Guardians' resistance and attacks Jack and Sandy. On the expedition, a young boy named Jamie Bennett (Dakota Goyo) is delighted to discover the Guardians in his room he believes in all of them except for Jack, whom he cannot see. Promising Jack that when they find Pitch, they will help him regain his own baby teeth, and through them the memories of his past life. To avert this, the Guardians travel the world collecting teeth,

Resentful that he himself is not believed in, Pitch plans to destroy children's faith in the Guardians and thereby weaken them by preventing Toothiana from doing her work. He also captures Toothiana's helper fairies, but a single fairy, Baby Tooth, is rescued by Jack. Meanwhile, Pitch and his nightmare minions raid Tooth's palace, taking all the children's teeth and the memories stored within. Jack, frustrated and hurt by centuries of isolation, declines the invitation, but North persuades him to cooperate by explaining their mission and the looming threat. Aster Bunnymund (Hugh Jackman), and the Sandman (Sandy for short) to the problem, North learns that Jack has been selected by the Man in the Moon as the new recruit. After alerting fellow Guardians the Tooth Fairy/Toothiana (Tooth for short) (Isla Fisher), the Easter Bunny/E. The Boogeyman (Jude Law), has returned and is threatening the children of the world. North, better known as Santa Claus (Alec Baldwin), becomes aware that Pitch Black,

The spirit of winter, Jack Frost (Chris Pine), is lifted with no previous memory from the depths of a frozen pond by the Man in the Moon, only to discover that no one can see, hear, or touch him.